Page 5 - Palombo_Ferretti _2005
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ARTICLE IN PRESS
                                M.R. Palombo, M.P. Ferretti / Quaternary International 126–128 (2005) 107–136  111

























































          Fig. 3. Molars of M. meridionalis from various Italian localities. (A–B) Upper Valdarno. (A) IGF 61, left M3; (B) IGF 87, right M3; (C–D)
          Pietrafitta (Farneta FU, late Early Pleistocene). (C) CET 79, right m3; (D) CET 384, right M2; (E–G) Crostolo (Farneta FU, late Early Pleistocene).
          (E) MSNRE-CRO3, right M3; (F) same specimen, lingual view; (G) MSNRE, left m3. All teeth, except F, are in occlusal view. Scale bar is 5 cm.


          Savignano mammoth to a species more primitive than   definitely less than in M. trogontherii and M. primigen-
          M. meridionalis.                                     ius. Last upper and lower molars (M3–m3; Figs. 3a
                                                               and b) possess from 11 to 14 plates (without talons), an
          2.1.2. Mammuthus meridionalis (Nesti 1825)           average enamel thickness of 3.2 mm, and a mean
            M. meridionalis is one of the best known Plio-     hypsodonty index (height/width) of 1.3 (Ferretti, 1998;
          Pleistocene elephants. The species is based on the rich  Lister, 1996a).
          material from the Late Pliocene/Early Pleistocene (ca.  The Florentine naturalist Giovanni Targioni Tozzetti
          2.0–1.75 Ma) Upper Valdarno (Valdarno Superiore)     (1775) was the first to describe remains of fossil elephant
          fluvio-lacustrine deposits. The skull of M. meridionalis  from various localities in Tuscany. Targioni Tozzetti
          is on the average higher than in earlier mammoth taxa,  referred all the specimens he studied to the Asian
          with a concave forehead and wide parietals (Fig. 2a).  elephant, E. maximus, thus not realizing the Italian
          Tusks are large, more massive and curved in males,   material pertained to a newly discovered extinct species.
          showing a double curvature (spiraling) generally more  Later, Nesti (1808), working on the abundant elephant
          accentuated than in Elephas and Loxodonta, but       remains from the Upper Valdarno, remarked the
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