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Animal Biodiversity and Conservation 39.2 (2016) 163
A
Italy
Tunisia
B Aeolian
Ustica Archipelago
Egadi Calabria
Archipelago
Pantelleria
Linosa Malta
Tunisia Lampedusa
Fig. 1. Study area.
Fig. 1. Zona de estudio
the Scarabaeoidea fauna of other possible source is distinctly Mediterranean and the main natural
areas: Sicily, Calabria, Tunisia, Malta, Lampedusa environments are maquis and garrigue landscapes
and the Egadi archipelago (Favignana, Marettimo and (Ronsisvalle, 1973; Baccetti et al., 1995; Pasta &
Levanzo). These possible source areas, which include La Mantia, 2003; Lo Cascio & Pasta, 2004; Nesos,
the nearest mainland areas and other non–volcanic 2013). Given the relative environmental homogeneity
islands in the region, were examined in two different of the considered islands, it is assumed that the simi�
scenarios (see 'Analysis and interpretation' heading). larity in richness and composition should be mainly
The main climatic and vegetation factors are reaso� related with the role played by dispersal processes
nably homogeneous among the considered volcanic (Cadotte, 2006).
islands (Agnesi & Federico, 1995; Cicala, 1997; Pasta
& La Mantia, 2003; Nesos, 2013). The climate of the Data source and systematic group
volcanic islands is typically Mediterranean (Agnesi &
Federico, 1995; Pasta & La Mantia, 2003; Blasi et al., The presence–absence data (see annex) were drawn
2005; Nesos, 2013). The vegetation, despite being from Arnone et al. (1995, 2001), Carpaneto et al.
altered to some extent by a long human presence, (2005), Agoglitta et al. (2006), Dellacasa & Dellaca�