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Animal Biodiversity and Conservation 39.2 (2016)                                163
                           A

                                                  Italy

                               Tunisia

B Aeolian

                               Ustica                    Archipelago

                               Egadi                                  Calabria
                               Archipelago

                               Pantelleria

                               Linosa                    Malta

                      Tunisia                     Lampedusa

Fig. 1. Study area.
Fig. 1. Zona de estudio

the Scarabaeoidea fauna of other possible source         is distinctly Mediterranean and the main natural
areas: Sicily, Calabria, Tunisia, Malta, Lampedusa       environments are maquis and garrigue landscapes
and the Egadi archipelago (Favignana, Marettimo and      (Ronsisvalle, 1973; Baccetti et al., 1995; Pasta &
Levanzo). These possible source areas, which include     La Mantia, 2003; Lo Cascio & Pasta, 2004; Nesos,
the nearest mainland areas and other non–volcanic        2013). Given the relative environmental homogeneity
islands in the region, were examined in two different    of the considered islands, it is assumed that the simi�
scenarios (see 'Analysis and interpretation' heading).   larity in richness and composition should be mainly
The main climatic and vegetation factors are reaso�      related with the role played by dispersal processes
nably homogeneous among the considered volcanic          (Cadotte, 2006).
islands (Agnesi & Federico, 1995; Cicala, 1997; Pasta
& La Mantia, 2003; Nesos, 2013). The climate of the      Data source and systematic group
volcanic islands is typically Mediterranean (Agnesi &
Federico, 1995; Pasta & La Mantia, 2003; Blasi et al.,   The presence–absence data (see annex) were drawn
2005; Nesos, 2013). The vegetation, despite being        from Arnone et al. (1995, 2001), Carpaneto et al.
altered to some extent by a long human presence,         (2005), Agoglitta et al. (2006), Dellacasa & Dellaca�
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