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366 FABIO LIBERTO ET ALII
A. Brancato collection, Syracuse, Italy (CB); S. Familia CLAUSILIIDAE J.E. Gray, 1855
Giglio collection, Cefalù, Italy (CG); Laboratory of Subfamilia ALOPIINAE A.J. Wagner, 1913
Cytogenetics and Molecular Biology, University of Tribus MEDORINII H. Nordsieck, 1997
Urbino, Italy (LCMBU); F. Liberto collection,
Cefalù, Italy (CL); Museo Naturalistico F. Minà Genus Muticaria Lindholm, 1925
Palumbo, Castelbuono, Italy (MNMP); Museo Type species: Clausilia scalaris L. Pfeiffer, 1850
Civico di Storia Naturale di Comiso (MSNC);
Museo Civico di Storia Naturale di Genova “G. Muticaria cyclopica n. sp. (Figs. 1–13)
Doria”, Italy (MSNG); Museo Regionale di Ter-
rasini (MRT); A. Reitano collection, Tremestieri EXAMINED MATERIAL. Holotype: Italy, Sicily,
Etneo, Italy (CR); I. Sparacio collection, Palermo, Siracusa, Epipoli, 37°05’20”N, 15°13’49”E, 122 m,
Italy (CS); R. Viviano collection, Palermo, Italy legit A. Reitano, 5.V.2015 (MSNC n. 4537). Para-
(CV). types: Siracusa, Epipoli, Castello di Eurialo,
37°05'20"N, 15°13'48"E, 112 m, legit A. Reitano
and A. Brancato, 8.XI.2012, 5 exx (LCMBU);
MATERIAL AND METHODS idem, 3 shells (MSNC n. 4537); idem, 49 shells (CL
n. 16514–16562); idem, 8.XI.2012, legit A. Re-
All specimens were collected by sight on the itano, 38 exx (CR); idem, 2 exx (MSNG), idem, 2
soil and under the rocks. Observations on ecology exx (MNMP); idem, 37°05'20"N, 15°13'49"E, 122
of these organisms were made directly in the field. m, legit A. Reitano, 5.V.2015, 6 exx, 13 shell (CL
Dry shells have been studied as regard size, colour, n. 16771–16789); idem, legit A. Reitano, 6.IV.2016,
morphology, sculpture, aperture, plicae and lamel- 8 exx (CL n. 16798-16805); idem, legit A. Reit-
lae, lunella and clausilium. In order to study and ano, 6.IV.2016, 28 exx (CS); idem, 37°05'20"N,
illustrate genital organs, the specimens were 15°13'48"E, 112 m, legit A. Reitano, 6.IV.2016, 3
drowned in water and fixed in 75% ethanol. Re- exx (MSNC n. 4538, 4539, 4540); idem, 8.VI.2016,
productive apparatus was extracted by means of 15 shells (CL n. 17293–177307).
scalpel, scissors and needles. Photographs were DESCRIPTION OF HOLOTYPE. Shell sinistral, di-
taken with a digital camera. Height and maximum mensions: height: 13.58 mm, maximum diameter:
diameter of the shell along with some parts of 4.2 mm, cylindrical-fusiform, decollate, rather ro-
genitalia were measured (in millimeters) by a bust, light yellowish-grey in colour; external sur-
digital gauge. Voucher specimens were stored in face with minute, raised, close ribs, 40 ribs on
collections listed above. Toponyms (place-names) penultimate whorl; last whorl with robust and spa-
are reported following the Portale Cartografico ced ribs; spire slowly and regularly growing, with
Nazionale (PCN, http://www.pcn.minambiente.it 4 whorls; last whorl tapering downwards, with el-
/PCN/), Map IGM 1:25000. Each locality and/or evated and curved cervical keel and lower basal
collection site is in the original language (Italian). keel; suture moderately deep; umbilicus closed;
All the specimens were studied by a Leica MZ square aperture, with 5 lamellae (on parietum and
7.5 steromicroscope. The taxonomic order and no- columellar side) and lunella and 5 plicae (on pal-
menclatural arrangement follow Nordsieck (2007, atum); on parietum, starting from suture, there are:
2013) and Bank (2011). very long parallel lamella, emerging in its anterior
portion and well prolonged inside the shell in its
posterior portion; short spiral lamella, deviating
RESULTS from centre of parietum to adhere to parallel
lamella; (upper) parietal tooth-like lamella; on
SYSTEMATICS columellar side there are a low columellar lamella
and an internal subcolumellar lamella; on palatum
Phylum MOLLUSCA Cuvier, 1795 (Fig. 3) there is an evident and raised lunella and,
Classis GASTROPODA Cuvier, 1795 starting from suture: two sutural plicae, the prin-
Ordo PULMONATA Cuvier in Blainville, 1814 cipal plica is robust in its posterior portion, whereas
Subordo STYLOMMATOPHORA A. Schmidt, 1855 its anterior portion, fused to anterior upper palatal