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Two news Clausiliidae (Gastropoda Pulmonata) of Sicily (Italy) 371

Siciliaria (Siciliaria) calcarae orlandoi n. ssp.
(Figs. 15–30)

EXAMINED MATERIAL. Holotype: Italy, Sicily,
Corleone, Rocca Busambra, Ficuzza, 27.IX.1981,
legit V.E. Orlando (MRT, n. 31040 Orlando col-
lection, written in the box and in the register:
Siciliaria calcarai n. subsp., det. H. Nordsieck);
Paratypes: same data of holotype, 4 exx (MRT, n.
31041/4 Orlando collection); Corleone, Bosco
Ficuzza, 25.IV.1971, legit V.E. Orlando, 2 exx
(MRT, n. 4903/4 Orlando collection); Monreale,
Ficuzza, Val di Conti, 23.III.1981, 2 exx (CS);
idem, legit I. Sparacio, 8 exx (CL n. 17276–
Figure 14. Geographic distribution of genus Muticaria in CE 17283); Monreale, Diga Scanzano, 31.XII.1989, 5
and SE Sicily (in yellow) with M. cyclopica n. sp. (triangle), exx (CS); Monreale, Bosco del Cappelliere,
M. brancatoi (star), M. syracusana (square) and M. neute-
boomi (dots). 2.I.1991, 21 exx (CS); idem, 28.XI.1993, 8 exx
(CS); Godrano, Rocca Busambra, Alpe Cucco,
21.II.2010, 5 exx (CS); Monreale, Bosco Ficuzza,
Genitalia with very short penial diverticulum, Ponte Arcera, 37°55’42” N; 13°23’01” E;
longer in M. neuteboomi and M. macrostoma mac- 27.IX.2009; 9 exx, (CL n. 5508–5516); Monreale,
rostoma, M. macrostoma scalaris and M. mac- Bosco del Cappelliere, Cozzo San Leopoldo,
rostoma oscitans. Only M. macrostoma mamotica 37°54’53”N, 13°22’57”E, 616 m, 2.IV.2016, 4
has a penial diverticulm similar to that of M. cyc- shells (CV); idem, 3 exx, legit R. Viviano (CL n.
lopica n. sp.; however, M. macrostoma mamotica 16446–16448).
has genitalia with a pleat on the internal wall of the OTHER EXAMINED MATERIAL. Siciliaria calcarae
penis (not present in M. cyclopica n. sp.) and a ven- calcarae (Philippi, 1844). Italy, Sicily, Palermo,
tricose shell (fusiform in M. cyclopica n. sp.) with San Ciro, 31.X.1986, 7 exx (CS); idem,
shorter parallel lamella and anterior portion of prin- 38°05’11”N, 13°23’07”E, 190 m, legit Sparacio I.,
cipal plica distinct from anterior palatal plica (fused 28.XI.2015, 2 exx (CL n. 16807–16829); Bagheria,
in M. cyclopica n. sp.). Monte Catalfano, 30.VI.2006, 28 exx (CS);
Preliminary molecular studies (Gregorini et al., Palermo, base Monte Grifone, Cimitero Santa
2008; Colomba et al., 2010; 2012) showed the ex- Maria di Gesù, 24.VIII.2014, 11 ex (CS); Fav-
istence of significant genetic differences between ignana Island, Grotta delle Uccerie, 37°57’04”N,
populations attributed either to M. syracusana, M. 12°18’18”E; 30 m, 11.IX.2010, 17 exx, 14 shells,
neuteboomi or M. brancatoi, including the topo- (CL n. 8414–8444); Calatafimi, Le Rocche,
typic ones. Moreover, further and more complete 37°54’14”N, 12°48’14”E, 493 m, 20.XI.2011, 6
molecular data (personal unpublished data), con- exx, 15 shells (CL n. 10763–10783); Scopello,
firmed these preliminary results; furthermore, by Torre Bennistra, 07.XII.2016, 3 exx (CR); Erice,
comparing cytochrome oxydase I (COI) partial se- Monte Castellazzo, 20.VI.2002, 26 exx (CR);
quences, specimens of M. cyclopica n. sp. turn out Castellammare del Golfo, Monte Inici, VI.1996, 5
to be genetically distant from all other Sicilian and exx (CR).
Maltese Muticaria populations. Siciliaria (S.) calcarae belliemi (Brandt, 1961).
Italy, Sicily, Partinico, Monte Belliemi, 1.III.2015,
28 exx (CS); idem, 8.V.2016, 34 exx (CS); idem 9
Tribus Delimini R. Brandt, 1956 shells (CL n. 17284–17292).
Genus Siciliaria Vest, 1867 Siciliaria (S.) ferrox (Brandt, 1961). Italy, Sicily,
Subgenus Siciliaria Vest, 1867 Trabia, Torre Sant’Onofrio, 143 m, 25.VIII.2007,
Type species: Clausilia grohmanniana Rössmassler, 30 shell (CL n. 2331–2360); Altavilla Milicia,
1836 Grotta Mazzamuto, 15.X.2015, 25 exx (CS).