Page 8 - Liberto_Reitano_et_allii_2016

372 FABIO LIBERTO ET ALII

DESCRIPTION OF HOLOTYPE. Dimensions: height the epiphallus; epiphallus (2.6–3.2 mm) divided by
19 mm; maximum diameter 4.8 mm. Shell elong- point insertion of robust penial retractor muscle into
ated, fusiform, sinistral, not decollate, obtuse apex, cylindrical-conic proximal portion and shorter
robust, brown in colour (Figs. 15, 16); external sur- distal portion slightly enlarged before entering in
face with very minute and just raised ribs equally the penis. Penis short (1.6–2.2 mm), wider than epi-
arranged in all whorl sof teleoconch; 92 ribs on pen- phallus; internal walls of penis show two weak lon-
ultimate whorl. Spire slowly and regularly growing, gitudinal furrows; conic penial papilla, with slightly
with 11 whorls little convex; basal and cervical pointed apex and a restriction to the base.
keels little distinct; umbilicus closed; suture shallow Body. Animal long, narrow, posteriorly pointed,
with papillae scattered and slightly evident (more blackish with a dorsal, narrow and whitish band;
numerous from third to seventh whorl); aperture skin tubercle ovale-elongated; upper tentacles
about ¼ of shell height, subovoidal, with 4 lamellae rather short, cylindro-conical, whitish, apically
on parietum and columellar side, lunella, and 4 widened with small black eyes; pneumostome and
plicae on palatum. On palatum there is an evident genital opening on left side; foot long, narrow, with
lateral lunella, starting from suture there are a long sole paler than body.
and raised principal plica not fused to lunella apex BIOLOGY AND DISTRIBUTION. Siciliaria calcarae
and slightly wider in its posterior portion, a short orlandoi n. ssp. lives under the bark of dead trees
posterior portion of upper palatal plica fused to lun- and in the leaf litter of woods vegetating both in
ella apex and an obsolete upper palatal plica re-
presented only by a short, large callosity little in re- sandstone (Bosco del Cappelliere, Diga Scanzano)
and calcareous (Alpe Cucco, Rocca Busambra)
lief, a medium long basal plica, the internal first soils (Figs. 29, 30); in these two last localities
part of which is joined to the base of lunella; a short
sulcalis. On parietum and columellar side there are: S. calcarae orlandoi n. ssp. is found also on cal-
careous rocks into the woods. This new subspecies
non emergent and well raised spiral lamella in is known for the “Nature Reserve Bosco della
centre of parietum; tooth-like (upper) parietal
lamella, moderately high (inferior) columellar Ficuzza, Rocca della Busambra, Bosco del Cappel-
liere e Gorgo del Drago” an area wich is included
lamella, non emergent subcolumellar lamella. in the Sicani Mountains Regional Natural Park
Peristome continous, slightly thickened, reflected,
superiorly attached to the wall of last whorl. since 2013.
Siciliaria calcarae s.l. lives on calcareous rocks,
VARIABILITY. Dimensions of paratypes (not de- in cavities and under stones on calcareous soils. It is
collate) (Fig. 17): height: 18–22 mm; maximum dia- described from Palermo and is widespread in West-
meter: 4.2–4.8 mm; ribs on the penultimate whorl ern Sicily and the Egadi Islands (see Beckmann,
of the shell ranges from 88 to 95 mm, but some ribs 2004) (Figs. 31, 32, 56).
are incomplete or obsolete; sometimes a very little ETYMOLOGY. The new subspecies is dedicated to
sutural plica is present; the upper palatal plica can
be very small or absent (Figs. 18, 19). Parietum as Vittorio Emanuele Orlando (1928–2014, Terrasini,
in figure 20. Clausilium (Figs. 21–22) with elongate Italy), who identified this taxon, to his passion for
molluscan studies and his museum activity in Sicily.
plough-like basal plate, sutural angle slightly bent
up, palatal and columellar edges of plate nearly par- COMPARATIVE NOTES. Siciliaria calcarae orlandoi
allel; outer corner more or less pointed. n. ssp. is distinct from S. calcarae calcarae (Figs.
Genitalia (5 specimens examined) (Figs. 23–28) 31, 35, 37–41, 45–55) for the reduced anterior
are characterized by: slender and thin free oviduct, upper palatal plica (longer and raised in S. calcarae
well developed ovispermiduct; bursa copulatrix calcarae who as, rarely, also a small second upper
complex consist of slender copulatory (3.45–2.8 palatal plica), reduced or absent sutural plica
mm) duct ending in two branches: one branch con- (present in S. calcarae calcarae), moderately high
sisting of a long diverticulum of the bursa copulat- columellar lamella (low in S. calcarae calcarae), the
rix (5.2 mm), second branch consisting of very clausilium with sutural angle slightly bent up, thus
short bursa copulatrix duct with cylindrical bursa palatal and columellar edges of plate are nearly
copulatrix; vagina short (1.8–2.5 mm) and uniform parallel (sutural angle much bent up in S. calcarae
in diameter; vas deferent long and slender, entering calcarae).