Page 3 - NAUTILUS_122-1_Garilli

V. Garilli, 2008
SYSTEMATICS
Family Rissoidae Gray, 1847
Subfamily Rissoinae Gray, 1847
Genus Alvania Risso, 1826
Type Species: Alvania europea Risso, 1826 [synonym
of A. cimex (Linnaeus, 1758)], subsequent designation by
Nevill, 1885
Alvania carinata (Da Costa, 1778)
(Figures 1–14)
Turbo carinatus Da Costa, 1778: 102–103, pl. 8, fig. 10
Rissoa trochlea Michaud, 1830: 16, fig. 4
Galeodina cingulata (Philippi, 1836).—Piani, 1979: 70–71, figs. tany, Finistère Roscoff, “les Cochons Noirs”, sand and
2–3
Description: Shell small, sturdy, conical and keeled to
slender and turrited, reaching about 5 mm (rarely 6 mm)
in height; 3.9–4.1 mm in width. Protoconch multispiral,
conical, consisting of about 2.2 convex whorls. Proto-
conch I with 0.8 whorls, sculptured with six very fine
spiral lirae and microscopic granules between them. Pro-
toconch/teleoconch transition well-marked and sinuous.
Protoconch II sculptured with sparse, spirally arranged
microscopic pimples, stronger in adapical direction, and
one to two spiral ridges, one of them always very close to
lower suture. In the largest shells, teleoconch formed by
4–4.5 moderately convex whorls. Common morph (Fig-
ures 2, 6–7) with teleoconch whorls markedly dominated
by spiral sculpture, which consists of very strong cords
(numbering 2–3, 3–6, 6–12, and 12–17 on first, second,
third, and last teleoconch whorl, respectively). Second-
ary, less conspicuous cords may occurr on last whorl.
More marked spiral cords on adapical portion of whorls,
at a certain distance from suture, give a characteristic
keeled shape. Cords progressively less strong on basal
area. Unkeeled morph (Figures 1, 4–5) characterized by
a slender shell shape, usually bears more spiral cords
(18–20) on last whorl. Axial sculpture always formed by
numerous (32–50 on penultimate whorl), occasionally
very narrow and lamella-like ribs, becoming obsolete to-
ward base. Intersection of ribs with spiral cords gives an
almost general clathrate pattern in unkeeled morph. In-
tersection of spiral and axial sculptures nodular, usually
forming squares (Figure 14), with exception of last whorl
where a rectangular pattern occurs (Figure 13). Micro-
sculpture consists of very fine spiral threads (Figures
13–14), covering all teleoconch surface, with exception of
main spiral pattern. On early teleoconch whorls, spiral
lirae often alternate with spiral alignments of micro-
scopic pimples. Sutures slightly inclined. Last whorl well
expanded, comprising 3/5 to 3/4 (rarely more than 3/4,
Figure 3) of shell height, often bearing one or two varices
(mainly in keeled morph) with angles of 10°–340°. Ap-
erture wide, ovate, slightly rounded to angled in the pos-
terior part, comprising 3/5 to 3/4 of last whorl height.
Outer lip slightly prosocline, internally smooth, exter-
nally markedly thickened by a strong rim very close to lip
edge, and covered by spiral cords. Inner lip moderately
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arcuate and rather thickened, with a very narrow to dis-
cretely expanded (Figure 3) callus delimiting a very small
umbilical chink.
Type Locality: Cornwall, southwestern England.
Material Examined: Great Britain: Cornwall, Fal-
mouth, 1 sh., coll. MF, 1974, E55A; Channel Islands,
Herm, 11 shs, coll. MF, 09.1974, E54A. Atlantic France:
Normandy, Carteret, 3 shs, coll. MF, Jul. 1973, E28A;
lower Normandy, St. Pair, 7 shs, MNHN coll. Denis,
1945; Brittany, Finistère Anse de Bertheaume, 20–30 m,
industrial dredging, 4 shs, MNHN coll. S. Gofas, 1978;
Brittany, Cote-du-Nord Plomanac’h, fissures of infralit-
toral rocks, 1 sh., MNHN coll. S. Gofas, 1973–78; Brit-
conchiferous gravel, 20 m, 27 shs, MNHN coll. Gofas,
Jul. 1994; Brittany, St. Lunaire, 20 shs, MNHN Coll.
Fischer, 1898; Brittany, St. Servan, 10 shs, MNHN coll.
Staadt, 1969; Brittany, St. Lunaire, 2 shs, MNHN coll.
Ph. Dautzenberg (figured in Bucquoy et al., 1884, pl. 35,
figs. 1, 2); St. Lunaire, 5 shs, MSNCS, 7173 and 7174, 20
Jun. 1970, on the beach at low tide; Brittany, Morlaix,
Saint Michel en Grève, 2 shs, coll. MF, 1976, E25A;
Brittany, Saint Jacut, 3 shs, coll. MF, 06.1975, E12A;
Brittany, Saint Jacut, 14 m, 6 shs, coll. MF, 04.1974,
E12B; Brittany, Carnac, Quiberon, 4 shs, coll. MF, 1970,
E16B; Brittany, Carnac, Quiberon, Pointe de Couquel, 2
shs, coll. MF, 1970, E11A; Brittany, Saint Malo, 7 shs,
coll. MF, 07.1973, E13A; Brittany, St. Malo, Lizardrieux,
1 sh., coll. MF, Aug. 1982, E69A. Atlantic Pyrenees,
Aquitaine, St. Jean de Luz, Cote Basque, infralittoral
rocks, 1 sh., MNHN coll. S. Gofas, 1980–81; Atlantic
Pyrenees, Aquitaine, St. Jean de Luz, 73 shs, MNHN
coll. H. Fischer, 1898; Aquitaine, Soulac, 1 sh., MNHN
coll. A. Dolfus; Aquitaine, Hendaye, 2 shs, coll. MF, Jul.
1976, E57A. Portugal: Algarve Sagres, Baie de Baleeira,
(37°00.7Ј N, 08°55.0Ј W), tide zone, 1 sh., MNHN, Mis-
sion Algarve, May 1988; Algarve Sagres, Ponta da Ba-
leeira, (37°00.3Ј N, 08°55.5Ј W), 17–23 m, 5 shs,
MNHN, Mission Algarve, May 1988; Algarve Sagres,
Pontal dos Corvos, (37°01.3Ј N, 08°58.3Ј W), at the foot
of falaise, 17–22 m, 5 shs, MNHN Mission Algarve, May
1988; Albufeira (southern coast), Ponta de Castelo, 3–6
m, 2 shs, coll. MF, 08 Aug. 1985, E50B. Atlantic Moroc-
co: Asilah, mouth of Oued el Helou, conchiferous depos-
its, beach, 6 shs, MNHN coll. S. Gofas, 1971–72. Strait of
Gibraltar: Tanger, Grande Plage, conchiferous deposits,
beach, 3 shs, MNHN coll. S. Gofas, 1970–81; Cadiz,
Getares, beach, 3 shs, coll. MF, ex coll. C. Bogi, legit
Hanselaar, 2230 GET; Cadiz, Barbate, conchiferous de-
posits, beach, 6 shs, MNHN coll. S. Gofas, 1976–81;
south Ceuta, Punta del Desnarigado, (35°53.6Ј N,
05°16.8Ј W), 16–20 m, 1 sh., MNHN coll. Bouchet, Go-
fas and Lozouet, May 1996. Mediterranean Spain: Cas-
tellon, Columbretes Islands, Espinosa Island, 5 m, 1 sh.,
coll. MF, 26 Jul. 1974, M273A; Malaga, 15 m, 7 shs, coll.
MF, ex coll. Cesare Bogi, 2127IMA(V002G); Malaga,
Algeciras, Torre del Almirante, 3–5 m, 1 sh., coll. MF, 28