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Carduus affinis var. brutius Fiori in Fiori & Bég., Fl. Italia 3(2): 356. 1904. [April 1904]
≡ Carduus brutius (Fiori) Huter, Oesterr. Bot. Z. 56(12): 479. 1906. ≡ Carduus affinis subsp. brutius (Fiori)
Kazmi, Mitt. Bot. Staatssamml. München 5(2): 423. 1964. ≡ Carduus affinis subsp. brutius (Fiori) Peruzzi,
Inform. Bot. Ital. 42: 530. 2010. isonym
Ind. Loc.:—“Cal. sett. sul Monte Mula a ca. 1850 m”. CAL
Type (lectotype, designated by Kazmi 1964: 423):—ITALY. Calabria III District. Castrovillari, in M.te Mula,
S. Donato (S. Sosti), loc. pinguib. dumetos. at arborum term., sol. calcar. 1850 m, 26 July 1877, Huter, Porta &
Rigo 616 (W).
Note:—The basionym’s author is attributed based on the family index at the beginning of the volume (Arts.
46.8 and 37.5 of ICN).
Compiled by:—F. Bartolucci, L. Bernardo, G. Galasso, L. Peruzzi

Carduus corymbosus Ten., Fl. Napol. 1(3): XLVIII. 1812.
Ind. Loc.:—none.
Type:—Not designated.
Note:—Tenore (1812) in the protologue did not quote any collection locality. He reported for the first time
generic localities in the Synopsis novarum plantarum, quae in Prodromo Florae Neapolitane (Tenore 1819):
“Habitat in incultis Principatus Ulterioris, Samnii et Aprutii”.
Compiled by:—F. Bartolucci, F. Conti

Carduus fasciculiflorus Viv., Fl. Cors. Prodr., App.: 6. 1825. [November 1825]
Ind. Loc.:—“H. Vignolla, et Parao in Corsica”. COR
Type (neotype designated by Arrigoni 2014: 194):—ITALY. e Sardinia, 1827, Moris s.n. (TO).
Compiled by:—A. Brunu, G. Brundu, I. Camarda

Carduus membranaceus Lojac., Fl. Sicul. 2(1): 168 (pl. 14). 1903.
Ind. Loc.:—“Bosco di Colobria, Verdura! Etna, Strada per Nicolosi (subtypicus)”. SIC
Type (lectotype designated by Aghababyan et al. 2008: 519):—ITALY. Verdura, May 1844, Gasparrini? s.n.
(PAL n. 10180, right-hand plant).
Compiled by:—G. Domina

Carduus nutans var. typicus f. perspinosus Fiori in Fiori & Bég., Fl. Italia 3(2): 352. 1904. [April 1904]
≡ Carduus nutans subsp. perspinosus (Fiori) Arènes, Mém. Mus. Natl. Hist. Nat. 24: 236. 1949. ≡ Carduus
micropterus subsp. perspinosus (Fiori) Kazmi, Mitt. Bot. Staatssamml. München 5: 337. 1964.
Ind. Loc.:—“nei luoghi aridi qua e là”.
Type:—Not designated.
Note:—The basionym’s author is attributed based on the family index at the beginning of the volume (Arts.
46.8 and 37.5 of ICN).
Compiled by:—F. Bartolucci, F. Conti

Carduus macrocephalus subsp. siculus Franco, Bot. J. Linn. Soc. 71(1): 48. 1975. [18 November 1975]
≡ Carduus nutans subsp. siculus (Franco) Greuter, Willdenowia 33: 52. 2003.
Ind. Loc.:—“Siciliae”. SIC
Type:—ITALY. In apricis herbosis montosis–Palermo alla Pizzuta, s.d., Todaro 421 (holotype, K).
Compiled by:—G. Domina

Carduus intermedius Lojac., Fl. Sicul. 2(1): 167(–168). 1903.
≡ Carduus pycnocephalus subsp. intermedius (Lojac.) Giardina & Raimondo, Bocconea 20: 10, 386. 2007.
Ind. Loc.:—“In luoghi erbosi calcarei aprici, Palermo a Castellaccio e nell’Isola di Pantelleria”. SIC
Type (lectotype designated by Aghababyan et al. 2008: 519):—ITALY. Monte Cuccio, 7 June 1828, Tineo
s.n. (PAL n. 10165).
Compiled by:—G. Domina

42 • Phytotaxa 196 (1) © 2015 Magnolia Press PERUZZI ET AL.