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492 N. MAIO, A. PETRACCIOLI, P. CROVATO, N. AMOR & G. ODIERNA

Trochoidea (Trochoidea) trochlea (Pfeiffer, 1846) posto”. Sacchi (1955a) reports finding specimens
of Helicella (Trochoidea) trochlea Drap. in Eastern
ORIGINAL DESCRIPTION. Pfeiffer, 1846, species Algeria at Capo Carbon, near Bugia and at Capo di
no. 302, p. 69, locus typicus: “Patria ?”: “Helix Guardia.
trochlea Pfr. - T. perforata, conico-turrita, trochlearis, In the L. Pfeiffer collection of the Senckenberg
costulato-striata, albida, interdum fusco-maculata Forschungsinstitut und Naturmuseum in Frankfurt
vel basi fusco-fasciata; anfr. 8 acute crenulato- am Main (Germany), there is a specimen reported
carinati, late exserti, utriuque planulati, ultimus as “Helix trochlea Pf., coll. O. Reinhardt ex Par-
basi subtilius et confertius striatus, convexiusculus, reyss” from "Sicilia" (h/d=0.81) and nine specimens
antice non descendens; apertura securiformis; perist. reported as “Trochula trochlea Pf., coll. C. Bout ex
acutum, margine basali arcuato, intus leviter labi- H. Rolle” from "Sierra Nevada, Hispania"
ato, columellari subdilatato, ad perforationem sub- (h/d=0.72). In the same collection there are other
reflexo. Diam. 7 2/3, alt. 8 mm [h/d=1.04] - Patria? specimens with labels indicating provenance from
(Coll. Cuming.)”. sites in Algeria and Tunisia (R. Janssen, pers. com.;
ExAMINED MATERIAL. The first faunistic re- see Table 3).
ports of T. trochlea date back to Morelet (1853,
sub Helix trochlea Pfr.) from the promontory of Trochoidea (Trochoidea) elegans (Gmelin, 1791)
Ippona, Capo di Guardia near Bona [present town
of Annaba, author’s note], Algeria, and to Bour- ORIGINAL DESCRIPTION. Gmelin, 1791, species
guignat (1864) who reports four varieties of He- no. 229, p. 3642, locus typicus “Barbaria et australi
licella trochlea Pfeiffer (var. minor, depressa, Europa”: “Helix elegans. H. testa pyramidali alba
fusco-maculata and hypozona) from Bugia [present umbilicata: anfractibus sex acutis planiusculis
town of Béjaïa, author’s note], Algiers and at marginatis. Habitat terrestris in Barbaria et australi
Capo di Guardia, near Annaba (Algeria). Bourguig- Europa. [Barbaria or Berberia: the Mediterranean
nat (1868) also reports it from Tunisia at Kamart coastal area of Barbary and the Barbary States:
(Tunis), in the ruins of Carthage and on the east- Tunisia, Algeria and Morocco, author’s note]”.
ern slope of Djebel Ahmar (Tunisia). Specimens ExAMINED MATERIAL. During the present
with this determination from Annaba are kept in study, some populations with particularly elevated
the Löbbecke Museum in Düsseldorf (1868, legit spire were found in the literature. Morphologically
Lischke) and have h/d ratios in the range 0.68- very similar to T. caroni, they were of uncertain tax-
1.04 (mean: 0.86) (Maio, pers. obs.; Figs. 29–31). onomic attribution. The reports are included to
Kobelt (1877) writes: “This very attractive species complete the information on this particular group
represents [=substitutes?] the group of the “Hel. of land snails.
Caroni” in Algeria, where it is found in the Prov- Sacchi (1955a) reports finding specimens of He-
inces of Algiers and Constantine in various licella elegans with elevated spire and morphologi-
points” (Fig. 32). De Saint-Simon (1882) reports cal characteristics very similar to H. elata on Ibiza.
specimens of Helix trochlea collected at Capo di In the same paper he reports “molto turricolate”
Guardia, near Bona, of which he examined the forms of H. elegans from Tunis and specimens of
radula. Letourneux & Bourguignat (1887) also re- Helicella (Trochoidea) elegans stated to belong to
port it from Tunisia, as well as from Bona, Bugia “forme assai alte di spira” from Southern France.
and Algiers (Algeria). Monterosato (1893) reports Sacchi (1954, 1955a) does not exclude the hypothe-
H. trochlea Pfeiff. from “Algeria a Bona e nel de- sis of the presence of turriculate forms in the Balea-
serto del Sahara (Mouss.)” and H. speciosa, ric Islands, though he considers them the result of
Monts. as “nuova forma vicina alla trochlea ma geographical isolation of populations of H. elegans
pur distinta” from Salaparuta (Trapani Province) (reported, moreover, by Bofill I Poch & D’Aguillar-
in Sicily (Prof. A. Palumbo). Amat (1924) as Helix (Trochula) elegans).
Wenz (1923) reports this species as a “Pliocen. Two historical reports of T. elegans by Ross-
S. - Diluv.” fossil at "Cap de Garde bei Bone” and mässler (1837, Tafel xxVI, Fig. 345, sub H. elegans
Joleaud (1936) also reports it as a fossil from Capo Gm.) and by Aradas & Maggiore (1840: specie
di Guardia in the “panchina quaternario antica del LxVI, Elice elegante H. elegans Gm) from Sicily