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              reconstructing local hydrological balance and vegetation cover. Shell δ O has
              been related to atmospheric conditions, and in particular to the oxygen isotope
              ratio of precipitation, relative humidity and temperature (Balakrishnan and Yapp
              2004). However, the imprinting of atmospheric conditions on the shell oxygen
              isotope ratio is not straightforward and it is  shown as  distinctive regional
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              relations (e.g. Zanchetta et al. 2005). Shell δ C is mainly a function of respired
              CO 2  from the diet, which for most of the species means vegetation (e.g. Stott
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              2002). Shell δ C thus reflects vegetation type (e.g. C3, C4), but also species-
              specific  feeding behaviour (calciophilous) and potentially the effect of
              environmental conditions on the plants carbon isotope fractionation.

              Here  we analyse  the  stable carbon and oxygen isotope of shells  from  several
              archaeological sites of the Iberian and Italian Peninsula and Sicily (Figure 1;
              Table 1). Through a comprehensive compilation of previously published data
              (n = 854)  we explore shell  isotope variations between the  Late Pleistocene
              (~14.5 to 10.7 ka cal BP) and the early-late Holocene (~10.7 to 2.5 ka cal BP)
              and discuss their implications to understand  millennia-scale changes in
              hydrological conditions and vegetation dynamics. This  work also aims at
              stimulating research efforts to complement the land snail archive  with other
              environmental proxies from multiple sites (speleothems, lake cores etc.).

                            Table 1. Archaeological sites reported in Figure 1

                  Site                        14 C age (ka cal BP)         Reference
                  1 - Los Castillejos         ~7.2 – 4.0      Yanes et al. 2011
                  2 - Marroquíes              ~4.1            Yanes et al. 2013a
                  3 - Arena de la Virgen and Casa Corona   ~8.5 – 12.2   Yanes et al. 2013b
                  4 - Bauma del Serrat del Pont   ~2.5 – 9.0   Colonese et al. 2013
                  5 - Grotta di Latronico 3   ~7.8 – 8.8      Colonese et al. 2010a
                  6 - Grotta del Romito       ~13.0 – 14.5    Colonese et al. 2007
                  7 - Grotta della Serratura   ~7.2 – 12.1    Colonese et al. 2010b
                  8 - Grotta d’Oriente        ~7.8 – 14.2     Colonese et al. 2011

              Materials and Methods

              Archaeological and modern shell values are compared at the regional scale. The
              comparison also includes modern shell isotope data from Italian Peninsula and
              Sicily (e.g.  Zanchetta  et al.  2005; Colonese  et al.  2014) not  associated  with
              archaeological sites (Figure  2A). Variations in shell isotope composition are
              also analysed on a site-by-site basis by correcting archaeological values for the
              modern  average  shell  isotope  data  at  each  site  (Δδ  =  archaeological-modern;
              Figure 2B).

              Results and discussion

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              Early to late Holocene shells in general are depleted in  O and  C compared to
              Late Pleistocene and modern counterparts. By contrast Late Pleistocene shells





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