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fall within the range of isotope values of modern specimens from the western
              Mediterranean  (Figure  2A).  Δδ  adjusts  values  for  site-specific environmental
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              condition (e.g. elevation, δ O of rainfall, temperature) and species-specific

















                  Figure 1. Study area showing the location of the archaeological sites (Table 1)

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              feeding  behaviour  (i.e.  δ C).  Late  Pleistocene  δ O and δ C are similar to
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              higher  than  modern  counterparts  (up  to  4‰  and  2‰  for  δ C  and  δ O
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              respectively), whilst Holocene δ O and δ C values are variably lower (up to
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              6‰ in both δ O and δ C). Δδ O and Δδ C values are positively moderately
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              correlated (r = 0.63; R  = 0.40; Figure 2B), implying a major mechanism that is
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              driving changes in both δ O and δ C values over time. Variations between dry
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              and wet conditions would explain the δ O and δ C values between the Late
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              Pleistocene and Holocene. Higher δ O and δ C values in the Late Pleistocene
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              generally  reflect  a  negative  water  balance,  which  in  turn  increases  the  δ O
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              values of rainfall (Bard et al. 2002) and the δ C of vegetation (e.g. Khon 2010).
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              However other factors are likely to be involved as δ O changes in the source of
              precipitation.


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               Figure 2. A) Shell δ O and δ C distribution in Late Pleistocene, early to late Holocene
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               and modern snails. B) Corrected shell δ O and δ C values between archaeological and
                 modern counterparts for each site. Δδ value of 0‰ means no difference between
                                  archaeological and modern shells.


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