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The marine limpet Patella ferruginea 271
(a)
1986
(b) 20
16
2003
12
8
4
0
0–1 1–2 2–3 3–4 4–5 5–6 6–7 7–8 8–9 9–10
FIG. 4 Size (measured on the longitudinal axis) frequency
distribution of P. ferruginea at Galéria, Corsica, in (a) 1985 (data
from Laborel-Deguen & Laborel, 1991b) and (b) 2009 (present
study). 2009
as souvenirs at the tourist site of Kerkouan, Carthage.
Accessibility is the most important factor influencing
collection, regardless of the legally protected status of the
species (Guerra-García et al., 2004; Espinosa et al., 2009a).
To protect threatened molluscs such as P. ferruginea we
recommend, based on our findings, use of marine protected
areas combined with effective control of human visitation,
as has been implemented on Asinara island and the Zembra
archipelago. Although exploitation is evident in several
areas of Sardinia (Cristo & Caronni, 2008), the marine
protected area of Asinara has the largest individuals and 0–1 1–2 2–3 3–4 4–5 5–6 6–7 7–8 8–9 9–10
highest densities in the Corsica–Sardinia region, and the
population in the Zembra marine protected area (estab- FIG. 5 Size (measured on the longitudinal axis) frequency
lished in 1977 and where human visitation is prohibited) distribution of P. ferruginea at Zembra in 1986 (adapted from
showed strong recruitment and had many large reproduc- Boudouresque & Laborel-Deguen, 1986), 2003 (adapted from
tive individuals, indicating a well-structured and viable Limam et al., 2004) and 2009 (this study).
population (Espinosa et al., 2009a). Changes in the popu-
lation on Zembra during the last 25 years have been positive
−1
(Fig. 5) in terms of density (from 0.7 m in 1986 to 2.65 m −1 (e.g. Algeciras Bay and Ceuta; Espinosa et al., 2009a), could
in 2009) and mean length (mean of specimens .2 cm was be a source of larvae for adjacent areas. Connectivity
4.4 cm in 1986 and 5.4 cm in 2009). Marine protected areas between populations needs to be guaranteed by establish-
(Asinara, Zembra: present study; Habibas and Chafarinas ment of a large number of small and proximate (no more
islands: Espinosa, 2009; Guallart et al., 2006) and also non- than 10–20 km apart) marine protected areas, as suggested
legally protected sites, many of them on artificial substrates by Boudouresque et al., (2005).
© 2013 Fauna & Flora International, Oryx, 48(2), 266–275
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