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is subject to secular and periodic fluctuations as well as irregular or random variations.

                  These variations have been shown to be  tightly correlated with the dynamics of the


                  Atmospheric Circulation Index (ACI) or  the ‘Vangengeim-Girs’ Index (Girs, 1974),

                  which characterises a dominant direction of air mass transport within Atlantic-Eurasian


                  region. Global climate changes, such as variations in atmospheric circulation and wind

                  stress, would affect the LOD (Klyashtorin, 1998). The LOD index is measured as the


                  difference between the actual (astronomical) length of the day and the standard day of

                  86400 SI seconds (Stephenson and Morrison, 1984). The longest available LOD time

                  series covers more than 350 years (McCarthy and Badcoc, 1986, Figure 3), but the time


                  series appears inconsistent (data prior to 1800 displayed little variation in comparison to

                  the 1800-1984 period, Figure 3). The International Time Bureau in Paris carries out


                  continuous registration of the LOD since 1800. These records corroborate the above

                  LOD index over the 1800-1984 period.


                        Air temperature has been recorded in  Europe since centuries and has been

                  therefore widely used to assess environmental changes on animal populations over a


                  long-term period (e.g. Gian-Reto et al., 2002). There is, however, no consistent time

                  series of sea surface temperature (SST) for the Mediterranean before the most recent


                  period. Some studies, however, showed that air temperature correlates quite well with

                  inshore sea temperature, so that we used air temperature as a proxy of SST (e.g.

                  Fromentin and Planque, 1996; Fromentin et al., 1998; Southward et al., 1988;


                  Beaugrand et al., 2002). We firstly focused on two time series of temperature of the

                  Northern Hemisphere (NH), because they were the longest ones available: (i) the annual


                  average of the NH air temperature estimated over the past 150 years from a grid box of

                  time series of land and ocean surface air temperature (Jones et al., 1997; 1999; 2001;


                  Parker et al., 1994, Figure 3) and (ii) an annual time series of NH temperature





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