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MONTESANTO ET AL.: DISTRIBUTION OF TYLOS IN THE MEDITERRANEAN 49
Table 1. List of the beaches sampled in the present study. WGS84 coordinates are reported. CCS, “Caruso Collection” samples; LIT, Data from Giordani-
Soika (1954); PSS, Present study samples; TE, Tylos europaeus; TP, Tylos ponticus.
No. Beach Area Latitude N Longitude E Data source Species
1 Alicudi Aeolian Is. 38°33 01 14°21 57 CCS TP
15°10 34 PSS TP
2 Brucoli Sicily 37°17 28 12°30 09 PSS TP
12°53 02 PSS TP
3 Calcara Sicily 37°59 25 12°39 46 PSS TP
14°40 05 CCS TP
4 Castellammare del Golfo Sicily 38°01 43 12°19 34 CCS TP
14°35 08 PPS TP
5 Cornino Sicily 38°05 38 12°51 38 CCS TE
13°37 10 CCS TE
6 Donnalucata Sicily 36°44 37 15°12 23 PSS TE
15°29 11 PSS TP
7 Favignana Egadi Is. 37°55 47 15°04 43 PSS TP
12°33 30 CCS TE
8 Filicudi Aeolian Is. 38°33 32 12°52 02 CCS TP
14°57 02 CCS, PSS TP
9 Foce Fiume Belice Sicily 37°34 58 14°47 25 PSS TP
14°34 14 CCS TP
10 Foce Fiume Naro Sicily 37°14 11 15°07 10 CCS, PSS TE
12°36 50 LIT TP
11 Fontane Bianche Sicily 36°57 42 13°19 42 LIT TE
15°07 23 PPS TE
12 Giampilieri Sicily 38°04 03 15°06 03 PSS TE
11°56 50 CCS TP
13 Isola delle Correnti Sicily 36°39 02 14°19 59 PSS TP
15°05 11 CCS TE
14 Lampedusa Pelagie Is. 35°30 46 15°30 03 LIT TP
12°54 50 CCS TE
15 Linosa Pelagie Is. 35°51 53 15°03 40 PSS TE
15°38 56 LIT TE
16 Lipari Aeolian Is. 38°26 30 14°17 09 PSS TE
15°23 37 CCS TP
17 Marina di Modica Sicily 36°42 20 12°30 32 CCS, PSS TP
14°18 04 PSS TP
18 Marina di Ragusa Sicily 36°46 47 12°44 40 PSS TP
13°02 56 CCS TP
19 Marzamemi Sicily 36°43 14 12°49 08 LIT, CCS TE
15°28 21 PPS TE
20 Mazara del Vallo Sicily 37°38 03 15°16 13 LIT, CCS TP
15°23 38 LIT TE
21 Mondello Sicily 38°12 00 12°38 42 LIT, PSS TP
13°09 41 PSS TP
22 Morghella Sicily 36°42 11 15°05 17 CCS TE
23 Oliveri Sicily 38°07 19
24 Pantelleria Pantelleria Is. 36°50 12
25 Paradise Bay Maltese Arc. 35°58 55
26 Playa di Catania Sicily 37°27 10
27 Ponte Schiavo Sicily 38°05 17
28 Portopalo, Menfi Sicily 37°34 39
29 Punta delle Formiche Sicily 36°39 44
30 Punta Faro Sicily 38°15 56
31 Ramla I-Hamra Maltese Arc. 36°03 42
32 Roccalumera Sicily 37°58 21
33 Ronciglio Sicily 38°00 04
34 San Blas Maltese Arc. 36°03 25
35 San Vito Lo Capo Sicily 38°10 33
36 Sciacca Sicily 37°30 10
37 Selinunte Sicily 37°34 57
38 Sindaro Marino Sicily 38°16 01
39 Siracusa Sicily 37°03 15
40 Spadafora Sicily 38°13 46
41 Torretta Granitola Sicily 37°35 07
42 Ustica Ustica Is. 38°42 29
43 Vaccarizzo Sicily 37°26 16
pecially in the south-eastern sector. The circum-Sicilian is- values being reported for the beaches at Playa di Catania
lands, with the exception of the island of Lampedusa, feature and Giampilieri, respectively. The grain-size results allowed
exclusively T. ponticus, whilst only T. europaeus and only T. the plotting of the box-plot shown in Fig. 3.
ponticus was recorded from the islands of Gozo and Malta
(Maltese archipelago), respectively. With respect to beach sediment particle mean diameter,
the two species appear to exhibit partial niche partitioning.
A list with the median particle diameter for a number of In fact, T. europaeus in general demonstrated a preference
Sicilian beaches from where Tylos were recorded is shown for finer particle sizes, being restricted to a range of particle
in Table 2. An analyses of such parameters reveals that the diameters extending from fine to coarse sand (according to
lowest median particle diameter value was reported for the the Wentworth scale), whilst T. ponticus could tolerate a
beach at Playa di Catania (0.344), whilst the highest such broader range of particle sizes, being recorded from beaches
value was reported for the beach at Giampilieri (1.771), characterized by fine to very coarse sand.
where the highest standard deviation value was also recorded
(1.201), whilst the lowest such value was recorded for the The Student’s t-test for the difference between the mean
beach at Ramla (0.099). A similar pattern for standard error distribution of the two targeted species (independent sam-
was reported, with the highest (0.118) and lowest (0.006) ples), using the median values of grain-size for each beach,
revealed an insignificant difference (t = 1.8431; df = 17;