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S. et C. Gofas, May 1991; Malaga, Calahonda, conchif- pania, Peninsula of Sorrento, Punta Penna, 2 shs, coll.
erous detritus, beach, 1 sh., MNHN coll.S. Gofas, 1976– medshells.com ex coll. G. Ruggieri, 1988; Italy, Sicily,
81; Malaga, Benalmádena, conchiferous detritus, beach, Palermo, 2 shs, MNHN coll. Locard; Sicily, 1 sh.,
6 shs, MNHN réc S. Gofas, 1991–93; Malaga, Mijas, MNHN coll. Petit, 1873; Palermo, 3 shs, ZMB ex coll.
detritus, 10 m, 4 shs, coll. Stefano Rufini; Malaga, Cabo Monterosato, 81013 (originally labelled by Monterosato
Pino, detritus, 10 m, 2 shs, coll. SR, (41.80g); Morocco, as Cingula tenera, 3, 1125, Palermo); Palermo, Arenella,
Cabo Negro, beach, 1 sh., MNHN coll. S. Gofas, det. W. 2 shs, ZMB ex coll. Monterosato, 81014, (originally la-
Ponder, 1986; France, Provence, Marseille, Curry, belled by Monterosato as “Cingula tenera var . . . , 2,
beach, 3 m, 2 shs, coll. PAL ex coll. C. Bogi, Jun. 1986, Arenella, Palermo”); Sicily, Siracusa, Capo Passero, 16
2037MAR; Provence, Marseille, La Baule, small beach at m, 3 shs, coll. medshells.com ex coll. G. Ruggieri, 05 Sep.
25 km west from Marseille, 9 shs, coll. PAL ex coll. C. 1987.
Bogi, Oct. 1986, 2035BAU; Provence, Le Dramont,
(43°24.7Ј N, 6°51.7 E), 22–30 m, 26 shs, MNHN réc. J. Habitat: This species is clearly limited to infralittoral
Pelorce, 1992; Provence, Les Embiez, cote Nord et Petit depths. In the upper part of its distribution, it seems to
Rouveau, (43°05Ј N, 5°47Ј E), rocks, algae, 0–1 m, 11 live in very shallow waters, on algae. It likely lives also in
shs, MNHN réc S. Gofas, Jun. 1995; Provence, the cavities occurring in infralittoral muddy sandy bot-
Marseille, Cap Morgiou, “calque de la Triperie,” toms.
(43°12.2Ј N, 05°26.9Ј E), muddy sand , inside cavity, 22
m, 4 shs, MNHN réc H. Zibrowius, Jun. 1996; Provence, Distribution: In the western and central Mediterra-
Marseille, Grand Congloue, (43°10.6Ј N, 05°24.2Ј E), 33 nean the species seems to be well distributed; Adriatic
m, 50 shs, MNHN réc. H. Zibrowius, Jun. 1996; and eastern Mediterranean occurrences should be veri-
Provence, Les Embiez, passe du Gaou, rocks, photophile fied. In the Atlantic it lives along the Moroccan coasts
algae, (43°04.3Ј N 5°47.4Ј E), 0–3 m, 1 sh., MNHN réc and in the Canary Islands. To my knowledge, there is no
S. Gofas, Jun. 1995; Provence, St. Clair, infralittoral fossil record of this species.
rocks, (43°08.2Ј N 6°23.2Ј E), 0–1 m, 1 sh., MNHN réc.
S. Gofas, Sep. 1992; Tunisia, Sfax, 4 shs, MNHN coll. Remarks: This small species is characterized by having
Staadt, 1969; Italy, Liguria, Portofino, 1 sh., coll. PAL, a variable shell shape and sculpture. The conical shells,
2038; Italy, Tuscany, Livorno, Antignano, 0.5 m, brown bearing a strongly cingulated sculpture, which markedly
algae on rocks, 1 sh., coll. PAL legit Bogi, Apr. 1999, prevails over the axial, are comparable with typical
2029; Livorno, under littoral rocks, 0.5–1.0 m, 5 shs, coll. keeled morph of Alvania carinata. In addition, Cingula
PAL ex coll. C. Bogi, 2039LIV; Livorno, Meloria, 10–30 species provided with a strong spiral sculpture are com-
m, 3 shs, coll. PAL ex coll. C. Bogi, 1995, 2031MEL; parable to A. tenera. The ovate, slender shells of this last
Livorno, Tuscan Archipelago, Island of Elba, Capoliveri, species, with a finely cingulated sculptural pattern, might
32 m, 2 shs, coll. PAL ex coll. C. Bogi, Aug. 1994, vaguely resemble some species of Setia H. and A. Adams,
2036CAPOL; Tuscan Archipelago, Gemini Island, 1854. As a consequence, Piani (1979) and Van Aartsen
(southern side of Island of Elba), 11 m, 3 shs, coll. PAL, (1982) included A. tenera in Galeodina, whereas Nords-
2034; Tuscan Archipelago, Isola del Giglio, Punta ieck (1968; 1972) placed the same species in Setia and
Fenaia, 32 m, 1sh., coll. medshells.com ex coll. G. Rug- Cingula respectively. The last two views should not be
gieri; Tuscany, Grosseto, Argentario, 25 m, 1 sh., coll. accepted. In fact, the species of Setia have a smooth or
medshells.com ex coll. G. Ruggieri, Jul. 1988; Italy, very slightly sculptured shell with more convex whorls
Lazio, Ostia, Tor Paterno, 33 m, 5 shs, coll. medshells. than A. tenera, and Cingula species are characterized by
com ex coll. G. Ruggieri; Lazio, Roma, Santa Marinella, shells usually lacking axial sculpture and having a very
ex reti, 2 shs, coll. medshells.com ex coll. G. Ruggieri; thick outer lip.
Italy, Sardinia, Oristano, Santa Caterina di Pittinurri, 5
m, 3 shs, coll. PAL ex coll. C. Bogi, 30 Aug. 1986, The shells from eastern Atlantic (Canary Islands and
2040SCP; Sardinia, Sassari, Island of Maddalena, beach, Morocco) do not show meaningful differences from the
3 shs coll. PAL ex coll. C. Bogi, 2028IMA; Sardinia, Mediterranean ones studied. With regard to the resem-
Nuoro, Sant’Antioco, Cala Lunga, 20 shs, coll. med- blance between A. tenera and A. carinata, it is manifest
shells.com ex coll. G. Ruggieri, 03.1989; Italy, Cam- in the keeled, conical-inflated shell shape shown by their
typical respective morphs, and in having a quite wide and
developed aperture. However A. tenera never shows va-

Figures 70–79. Alvania rosariae new species 70. Protoconch of the holotype, lower Pleistocene of Cartiera Mulino, bed 3D1 of
Costa (1989), SE Sicily, Ragusa, Vittoria. 71. Protoconch of paratype 16, showing variation of the abapical ridges on protoconch II,
same locality and bed. 72. Dorsal view of protoconch of paratype 21, showing protoconch I sculpture and the abapical ridges on
protoconch II, middle to upper Pleistocene of Kyllini, NW Peloponnesus, N2 bed of Garilli et al. (2005a). 73–74. Holotype, detail
of early protoconch (73) showing sculpture of protoconch I and protoconch I/protoconch II boundary, and dorsal view of protoconch
(74). 75. Sculpture of protoconch I, paratype 21. 76–79. Holotype, detail of teleoconch sculpture: early whorls (76), first to second
whorl (77), showing microsculpture, penultimate to last whorl (78), showing the microscopic incremental scars, and last whorl (79).
Scale bars: 100 ␮m in Figures 70–72, 74, 76 and 78–79; 50 ␮m in Figure 77; 20 ␮m in Figure 75. Black and white arrows indicate
the protoconch I/protoconch II and protoconch/teleoconch boundaries, respectively.