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Ind. Loc.:—“In aridis alpinis montium Aprutii; Monte Costone, Monte Intermesole presso Aquila”. ABR
Type (lectotype designated by Del Guacchio & Caputo 2005: 571):—ITALY. In Italiae montibus, Monte
Corno presso Aquila, August 1823, Gussone s.n. (NAP).
Compiled by:—A. Santangelo, A. Stinca

Asperula deficiens Viv., Fl. Cors. Prodr., App. Alt.: 3. 1830. [15 March 1830]
≡ Asperula tomentosa var. deficiens (Viv.) Nyman, Consp. Fl. Eur. Suppl. 2: 156. 1889.
Ind. Loc.:—“H. in parva insula Tavolara dicta, Corsicae attigua”. SAR
Type (lectotype designated by Diana Corrias 1980: 289):—ITALY. Asperula deficiens Viv.! / ex Herbario
Viviani (TO).
Compiled by:—A. Brunu, I. Camarda

Asperula garganica Huter ex Ehrend. & Krendl, Bot. J. Linn. Soc. 68(4): 268. 1974. [30 October 1974]
Ind. Loc.:—“Gargano, in rupestribus elatioribus montis St. Angelo”. PUG
Type:—ITALY. Italia australis, Apulia: Gargano, in rupestribus elatioribus montis S. Angelo,
2000–2200ˈ, 23 June 1875, Porta et Rigo 579 (holotype, W).
Note:—Ehrendorfer & Krendl (1974) originally attributed the name of this species to Huter, Porta & Rigo,
referring to the collection n. 579 of these authors. However, on the original label, only Huter is quoted. Hence, only
the latter author, followed by “ex”, can be inserted before the names of the publishing authors.
Compiled by:—M. Villani

Asperula gussonei Boiss., Diagn. Pl. Orient. s. 1, 10: 63. 1849. [March–April 1849]
≡ Asperula cynanchica var. gussonei (Boiss.) Fiori, Nuov. Fl. Italia 2(4): 503. 1927.
Ind. Loc.:—“Hab. in rupibus excelsioribus Nebrodum Siciliae”. SIC
Type (lectotype designated by Brullo et al. 2009a: 86):—ITALY. Madonie, all’ovest del Monte Scalone, 30
June, Gussone s.n. (NAP).
Note:—According to Art. 60.7 of ICN the spelling of the specific epithet originally published as “Gussonii” is
to be corrected in “gussonei”.
Compiled by:—C. Salmeri, S. Brullo

Asperula cynanchica var. lactea Huter, Porta & Rigo ex Galasso var. nov.

It is distinct from A. cynanchica for the constantly milky-white flowers, from Asperula aristata sensu lato for the
flowers with tube ± as long as perianth lobes (G. Galasso).

≡ Asperula montana var. lactea Huter, Porta & Rigo, Sched. Itin. Ital. 3: 253. 1878. [after 8 January 1878] nom.
nud. ≡ Asperula lactea Brullo, Gargano, N.G.Passal. & Peruzzi, Inform. Bot. Ital. 42(2): 529. 2010. nom. nud. ≡
Asperula lactea (Huter, Porta & Rigo ex Galasso) Brullo, Gargano, N.G.Passal. & Peruzzi comb. nov. Bas.:
Asperula cynanchica var. lactea Huter, Porta & Rigo ex Galasso
Ind. Loc.:—“Calabria II orient. in rupibus calcar., perpendicolaribus m.te Consolino ad Stilo 5–700 m”. CAL
Type:—ITALY. Calabria II orient. in rupibus calcar., perpendicolaribus m.te Consolino ad Stilo 5–700 m, 14
June 1877, Huter, Porta et Rigo 255 (holotype, FI).
Note:—The name published along with Huter, Porta & Rigo’s collection n. 255 is actually a nomen nudum,
which is validly described and combined at species level here. We choose here to describe a variety and
simultaneously propose a combination (made by different authors) to maintain almost the same authorsphip used
up to now for this species. This is allowed by ICN (see also Crouch et al. 2014).
Compiled by:—G. Galasso, L. Bernardo, N.G. Passalacqua, L. Peruzzi

Asperula peloritana C.Brullo, Brullo, Giusso & Scuderi, Anales Jard. Bot. Madrid 66(1): 86(–90, 92, figs. 2,
3b, 6, tab. 1, map). 2009. [6 July 2009]
Ind. Loc.:—“Sicily, Peloritani, Monte Scuderi, lungo il versante settentrionale”. SIC

24 • Phytotaxa 196 (1) © 2015 Magnolia Press PERUZZI ET AL.