Page 21 - CAPPARIS_2006

142 Annals of the
Missouri Botanical Garden

Ghorashi-Al-Hosseni 495G (RGN). KAZAKHSTAN. Tchavto- Shrub procumbent; twigs straight, semi-erect,
gorozi Ioshnee sel. Tamerlanovski, Priajin s.n. (K). MON- sometimes reaching up to 3 m long, glaucous;
GOLIA. Tian-Shan, Mongolia, Potanin s.n. (K); Eastern internodes 1.5–5 cm; stipules curved, retrorse, not
Mongolia, Gobi, Prezwalski s.n. (K). TURKEY. Artvin,
Coruh, Davis & Hedge D32427 (E); Hakkari, Kalolans, Davis decurrent or somewhat decurrent, golden yellow, 0.3–
23870 (E); Kagizman, Watson 375 (K); Karabu ¨k, Baytop 0.6 cm long, 0.1–0.3 cm wide at the base. Leaves
11383/65902 (E); Kars, Davis 46680 (E) (K); Konya, elliptic to rounded, sometimes ovate, 2.5–4.5 3 2–
Akyokus, Dural 576 (E); Osmaniye, Balls 1199 (K); Sinop, 3.5 cm, herbaceous; indument very dense, whitish,
Kargi, Tobey 2809 (E); Tokat-Niksar, Davis 24882 (K);
Zonguldak, Karabuk, Davis & Coode D39050 (E). TURK- trichomes thin and long, 20–25 3 200–500 mm; leaf
MENISTAN. Aschabad, Freyn 260b (E); Syr-Darja, Golike veins not prominent; bases rounded, apices obtuse or
s.n. (K). UKRAINE. Krym, Yalta, Toige 1896 (RNG). acute; mucro long, 1–1.5 mm, straight or somewhat
UZBEKISTAN. Altyn Tepe, Tashkent, 490 m, Vasak s.n. curved; petioles short, 0.5–1 cm. Flower buds round-
(K); Chauvast, Samarkand, Paulsen 277 (C).
ed or acute; floral pedicels thick and short, 2.5–
3.5 cm; flowers zygomorphic; abaxial (odd) sepal
There is a clear reference in the protologue to the galeate, 1.5–2.2 cm long, 0.7–1.1 cm deep; stamens
Caucasus, which indirectly points to the specimens 100 to 150, anthers 3.5–4 mm, with acute apices.
from this area in the herbarium Willdenow (surpris- Fruit oblong, pulp red; ripe seeds dark brown, 2.7–3
ingly the protologue does not mention the Mussin- 3 2.6–2.8 3 1.8–2 mm.
Pushkin expedition, vid. infra.). The reference in the
protologue to a ‘‘Capparis herbacea. Marschall Illustrations. Mandaville (1990: pl. 69–70).
a Bieberstein’’ is presumably pointing to Capparis Phenology. Flowering from May to September.
ovata M. Bieb (Bieberstein, 1808, 1819). In the Distribution and habitat. Irano-Turanian and
herbarium Willdenow (B) are two sheets within folder Saharo-Arabian Regions. North Africa, Middle East
nr. 10034 pertaining to Capparis herbacea (Hiepks, into Pakistan [Afghanistan, Iran, Iraq, Israel,
1972). The folder is labelled: ‘‘Polyandria Monogynia/ Pakistan, Saudi Arabia, Yemen]; also in Egypt
Capparis herbacea foliiis/ subrotunde ellipticis ovalis/ (Zohary, 1960). Oasis in semi-deserts, flooding
axillis spinosis. Pedunculis/ unifloris/ Habitat Cauca- plains, sometimes in somewhat saline soils; at
sus.’’ Sheets 1 and 2 are numbered and annotated both elevations from 0 to 1000 m.
as follows: ‘‘C. herbacea’’ and ‘‘W.’’ The folder has two The sheet with the lectotype of Capparis leucophylla
other labels: ‘‘Mussin Pushkin/ W.’’ This refers to is labelled: I ‘‘Capparis leucophylla DC/ (De Candole
Count Apollo Apollosovich Mussin-Pushkin (1760– Script.)’’ II ‘‘de Bagad a ` Alep./ Olivier de Bruguiere’’
1805), who was a Russian explorer and plant III ‘‘Herb. Mus. Paris./ Capparis leucophylla DC.’’ The
collector. He led a botanical expedition to the coincidence between labels and references to type
Caucasus in 1800–1802. The other label is almost material in the protologue led us to suppose Candolle
illegible. It is not clear whether the specimen was was implicitly designating a holotype, provided the
originally collected in the Caucasus by Mussin- author used only one element.
Pushkin or, presumably, cultivated in the Royal
Selected specimens examined. AFGHANISTAN. Baghlan,
Botanic Garden of Berlin from seeds gathered by this
SW of Doshi, Hewer 1153 (K); Morghak, Furse 7735 (K).
collector (the species is published in a catalog of IRAN. Bam, Leonard 6052 (K); Kerman, Parris 75405 (E);
plants actually grown at Berlin). This species was, in Rudak, [5 Dehbarek], Davis & Bokhari D. 56505 (E). IRAQ.
fact, cultivated in another botanic garden in Saint Hamah, Karin & Noori 39981 (K); Tell Kotchek-Senonal,
Petersburg: ‘‘Cultam in tepidario saepe fruticosam Mosul-Liwa, J. B. Gillet 10846 (K); Southern Desert, S. of Al
Salman, Ravi, Agnew & Haines 1656 (E); Shaikhiya, Salma-
fieri et C. spinosae assimilari nunciat Fischer in
Samawa, Al-Shehbaz s.n. (RNG). ISRAEL. Gilboa mountain,
litteris’’ [Friedrich Ernst Ludwig von Fischer was Davis 4668 (E). JORDAN. Jordan M.E.D.P. S/1113 (K).
director of the Imperial Botanic Garden of Saint PAKISTAN. Chitral, Tirich, Stainton 2780 (E). SAUDI
Petersburg (Pritzel, 1872)] (Bieberstein, 1819). ARABIA. Rumah, White 73 (K). YEMEN. Huth, Miller
3156 (E); Sanaa, Miller 3401 (E).
8c. Capparis sicula Veill. subsp. leucophylla (DC.)
8d. Capparis sicula Veill. subsp. mesopotamica
Inocencio, D. Rivera, Obo ´n & Alcaraz, stat. et
Inocencio, D. Rivera, Obo ´n & Alcaraz, subsp.
comb. nov. Basionym: Capparis leucophylla DC.,
nov. TYPE: [Iraq] ‘‘Amara, 9 Sep. 1918, W.
Prodr. Vol. I: 246. 1824. TYPE: [Iraq] ‘‘Inter.
Edgar Evans, M/100 (E)’’ (holotype, designated
Bagdad et Alep. Oliv. et Brug. (v.s. in Herb. Mus here, E!, specimen E 65908). Figure 7.
Par.)’’ (lectotype, designated here, P!).
Indumentum trichomatibus brevioribus, 100–250 mm
Capparis spinosa L. var. pubescens Zohary, Bull. Res. Council longis, et tenuioribus, 15–20 mm latis, internodiis aculeis
Israel 8D: 56. 1960. TYPE: [Egypt] Galala, rocks, ad basim tenuioribus, 0.1–0.2 cm, pedicelis brevioribus, 3–
4100 ft., 1944, P. H. Davis 8062 (holotype, K!). 4.5 cm, a typo differt.