Page 19 - CAPPARIS_2006

140 Annals of the
Missouri Botanical Garden

4b. Leaf almost glabrous; stipules somewhat curved, Valimiu ´, Peloponessos, Hanson 67–1-1 (RNG). ITALY.
retrorse or spreading, golden yellow contrasting Basilicata, Potenza, Akeroyd 3328 (RNG); Gigerti, Agrigento,
with the light color of twigs; anthers very small, Tavalione s.n. (C). MOROCCO. Fez, Inocencio 60016 (MUB);
1.6–2 mm ....... .. 8e. C. sicula subsp. sindiana Khe ´niche `t, Inocencio 60014 (MUB); Mekne `s, Jury 15502
2b. Leaves appearing white due to dense pubescence (RNG); Sidi-Karcen, Inocencio 60017 (MUB). SPAIN.
..... .......... 8c. C. sicula subsp. leucophylla Alicante: Reservoir of Amadoiro, Alcaraz 48705 (MUB);
La Alguen ˜a, Crevillente, Inocencio & Alcaraz 48683 (MUB).
Distribution. Figure 5. Almerı´a: Santa Marı´a de Nieva, David 1017 (RNG); Nı´jar,
Inocencio & Alcaraz 48712 (MUB); Berja, Inocencio &
8a. Capparis sicula subsp. sicula Alcaraz 48710 (MUB); Can ˜ada de Ve ´lez, Orce, Inocencio &
Alcaraz 48714 (MUB); Las Cuadras de D. Pepe, Velez Rubio,
Capparis spinosa var. canescens Coss., Anal. Sci. Nat., 11: 28. Inocencio & Alcaraz 48682 (MUB); Los Lobos, Inocencio &
1849. Capparis spinosa subsp. canescens (Coss.) A. & Alcaraz 48723 (MUB); Church of Saliente, Chirivel, Inocencio
O. Bolo `s, Misc. Fontsere: 88. 1961. Capparis ovata & Alcaraz 48713 (MUB); Hue ´rcal-Overa, Inocencio & Alcaraz
Desf. var. canescens (Coss.) Heywood, Feddes Repert. 48728 (MUB); Sero ´n, Inocencio & Alcaraz 48678 (MUB);
69: 56. 1964. TYPE: [Spain] ‘‘In rupestribus oppiduli Sierra de Gador, Valde ´s 813 (RNG). Ca ´diz: Jerez, Inocencio
Buenavista prope Xerez (E. Bourgeau, pl. Esp. 1849. & Alcaraz s.n. (MUB); Sanlu ´car, S. Silvestre 2524/68 (RNG).
n. 43)’’ (lectotype, designated here, P!). Granada: Castillejar, P. F. Cannon 692 (RNG); Mala ´,
Capparis ovata var. palaestina Zohary, Bull. Res. Council Gardner 1337 (RNG); Lanjaro ´n, Inocencio & Alcaraz 48718
Israel 8D: 55. 1960. TYPE: [Israel] Upper Galilee, (MUB); Reservoir of Negratı´n, Inocencio & Alcaraz 48703
Wadi Hindaj, 25 June 1954, M. Zohary 110 (holotype, (MUB); Guadix, Inocencio & Alcaraz 48712 (MUB); Loja,
HUJ!). Inocencio & Alcaraz 48681 (MUB); Baza, Inocencio & Alcaraz
48680 (MUB); Salobren ˜a, Inocencio & Alcaraz 48717 (MUB).
Shrub procumbent; twigs straight, sometimes reach- Huesca: Velilla de Cinca, Inocencio & Alcaraz 48688
(MUB). Ma ´laga: Torre del Mar, Inocencio & Alcaraz 48688
ing up to 3 m long, dark green; internodes 1.5–5 cm;
(MUB); Velez Ma ´laga, Inocencio & Alcaraz 48719 (MUB).
stipules curved, retrorse, not decurrent, occasionally Mallorca: from Palma de Mallorca to Andraitx, Inocencio &
slightly decurrent, yellow-orange, 0.3–0.6 cm long, Alcaraz 48726 (MUB). Murcia: Santa Cruz del Calvario,
0.2–0.3 cm wide at the base. Leaves ovate, oblong or Lange s.n. (C); Murcia, Lange s.n. (C); Venta del Olivo,
elliptic, 3–5 3 2–4.5 cm, herbaceous; indument lax, Inocencio & Alcaraz 48708 (MUB); Barinas, Inocencio &
Alcaraz 48704 (MUB); Albudeite, Inocencio & Alcaraz 48709
trichomes thick and long, 20–50 3 400–800 mm; leaf
(MUB); Lorca, Inocencio & Alcaraz 48722 (MUB); Ye ´char,
veins not prominent; base rounded or somewhat Inocencio & Alcaraz 48711 (MUB). Jae ´n: Alcaudete,
tapering, apices acute; mucro long, 1–1.5 mm, usually Inocencio & Alcaraz 48721 (MUB); from Larva to Guadiana
curved; petioles short, 1–1.5 cm. Flower buds acute; Menor, Inocencio & Alcaraz 48727 (MUB); Jo ´dar, Inocencio &
Alcaraz 48720 (MUB). Sevilla: Osuna, Inocencio & Alcaraz
floral pedicels thick and long, 5.5–7 cm; flowers
48679 (MUB); S of Spain, J. W. Carr 20706.6 (RNG).
zygomorphic; abaxial (odd) sepal galeate, 1.7–2.5 cm SYRIA. Djebel Casioun, Damascus, Sammuelsson 1518 (K).
long, 0.7–1.2 cm deep; stamens 100 to 150, anthers TURKEY. Ankara, Davis & Coode 37217 (E); Dardanelles,
2.5–3 mm, with acute apices. Fruit oblong, pulp red; s.n. (E); Denizli, K. Hormia 588 (RNG); Erzurum, Oltu,
ripe seeds dark brown, 2.6–3 3 2.4–2.8 3 2–2.2 mm. Baytop 14366 (E); Hakkari, Zab river, Trelawny 1305 (E);
Keslitu ¨rkmenli, Mersin, Hennipman 1073 (RNG); Pamuk-
Illustrations. Boccone (1674: 79, tab. 42. f. 3); kale, Denizli, Baytop 10231 (E); Saraycik-Osmancik, Tobey
2569 (E).
Zohary (1966: 359); Valde ´s et al. (1987: 374);
Castroviejo et al. (1993: 520, pl. 142); Plitmann et 8b. Capparis sicula subsp. herbacea (Willd.)
al. (1983: 81); O. Fragman et al. (2001: 312–3, Inocencio, D. Rivera, Obo ´n & Alcaraz, stat. et
pl. 133). comb. nov. Basionym: Capparis herbacea Willd.,
Phenology. Flowering May to October. Enum. Pl.: 560. 1809. Capparis ovata var.
Distribution and habitat. Mediterranean and herbacea (Willd.) Zohary, Bull. Res. Council
Irano-Turanian Regions; locally introduced in the Israel 8D: 56. 1960. TYPE: [Russia, Georgia or
Saharo-Arabian Region. Mediterranean Europe, North Azerbaijan] ‘‘Capparis herbacea. Marschall a Bie-
Africa, Middle East into Turkey [Albania, Algeria, berstein. Habitat ad Caucasum’’ (lectotype,
Cyprus, Greece, Italy, Morocco, Spain, Syria, Turkey]. designated here, B!, unlabelled blossoming
Stony places, marls or clayish soils, scrubs, at specimen, B-W, folder 10034, right of sheet 2).
elevations from 0 to 600 m; in the vicinity of human
dwellings. Capparis herbacea Willd. var. microphylla Ledeb., Fl. Ross.:
235. 1842. Capparis leucophylla DC. var. microphylla
Selected specimens examined. ALBANIA. Below Levani (Lebed.) Ta ¨ckh. Students’ Flora of Egypt, Ed. 2: 164.
and Frascula district, M. Barat, Baldaci 197 (K). ALGERIA. 1974. TYPE: [Kazakhstan or Turkmenistan] ‘‘Hab. ad
Ora ´n, Faure s.n. (K). CYPRUS. Cape Greco, Gold 29 (RNG). litt. Orient.m. Caspii! Karelin’’ (holotype, LE not seen).
GREECE. Baths of Eretria, Island Euboea, Stamatiadhou
17290 (C); Crete, Kissamos, Bickerich 15091 (K); Irakliou, Shrub procumbent; twigs straight, sometimes reach-
Bowen 8913 (RNG); Fokis, Delfi, Pinset 107 (RNG); Island ing up to 3 m long, light green, herbaceous;
Poros, Strid 29680 (C); Mitilini, Lesbos, Hansen 4918 (C); internodes 1–3 cm; stipules straight, spreading, not