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816 V. FIORENTINO ET AL.

Figure 6. A, geographic distribution of populations and lineage patterns as revealed by phylogenetic analyses. Black
pentagons indicate distribution of ‘Marmorana muralis’. B, consensus tree (50% majority rules) from Bayesian analysis
based on combined cytochrome oxidase subunit I and 16S sequence data (967 bp). Posterior clades probability and
maximum parsimony bootstrap values > 70% of clades are reported at nodes. Outgroup: Marmorana serpentina (Si,
Siena). For abbreviations, see Appendix 1.

the five species recognized in the past on the basis 2008), shell shape appears to be a variable character
of shell characters were not supported by exclusive in the Marmorana group.
mitochondrial lineages. Clades containing popula- Shell differentiation is faster than that of mitochon-
tions with globular/depressed and flat keeled shapes drial genes (Haase et al., 2003) and, moreover, intro-
also included intermediate shapes. Because appar- gression of haplotypes by hybridization between
ently no macro-environmental data were correlated populations with globular and flat shells has been
with Marmorana shell shapes (Fiorentino et al., hypothesized as a secondary source of shell variability

© 2008 The Linnean Society of London, Biological Journal of the Linnean Society, 2008, 94, 809–823

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