Page 10 - CAPPARIS_2006

Volume 93, Number 1 Inocencio et al. 131
2006 Revision of Capparis Sect. Capparis

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Figure 2. Distribution map for Capparis ovata ( ) (both subspecies); Capparis spinosa ( ); and Capparis aegyptia (X).
*
Capparis aegyptia also occurs in India, beyond map range.
oblong, pulp color unknown; ripe seeds dark brown, Selected specimens examined. EGYPT. Cairo, Schwein-
3.2–3.4 3 2.8–3 3 1.8–2 mm. furth 995 (K); Gebel Ez Zebir, Sinai, Tadmor S-417 (K);
Minia, Lippi ? (P); Tadmor & Shmida S-420 (E); Sinai, Bove ´
273 (K). INDIA. Mahablesham, Bombay, T. C. s.n. (E).
Illustrations. Plate 31(3) in Delile, 1812. Zohary
ISRAEL. Nabulus, Stait 266 (RNG); Wadi Qelt, Davis 4888
(1960: 52, fig. 1). Ta ¨ckholm. (1974: 163, pl. 48c). (K); Ein Gedi, Lyschede s.n. (C); Kfar Gileh and Manara,
Migahid (1988: 48, pl. 26). Curle 65 (K); Manara, Curle 143 (E); Mount Gilboa, Davis
Phenology. Flowering and fruiting from (January) 4667 (E); Wadi Yarmuk, Davis 4604 (E). JORDAN. Wadi
March to August (December). Mujib, Ma’daba, Boulos 5856 (K). SAUDI ARABIA. Wadi
Lakus, Jebel, Collenette 7228 (K).
Distribution and habitat. Saharo-Arabian, extend-
ing into the Irano-Turanian and Mediterranean
Regions. North Africa, Middle East into India 2. Capparis atlantica Inocencio, D. Rivera, Obo ´n&
[Egypt, India, Israel, Jordan, Saudi Arabia]. Rocky Alcaraz, sp. nov. TYPE: [Morocco] ‘‘Safi, 20–6-
places, steep slopes, at elevations from 0 to 2000 m, 1999. Inocencio 60026’’ (holotype, UMH!; iso-
often in the vicinity of human dwellings. It is the types, MO!, K!, E!, MA!). Figure 3.
common caper in Egypt and is often associated with Suffrutex decumbens, usque ad 60 cm, caulibus purpure-
the Hyparrhenia hirta (L.) Stapf community. Figure 2. is, foliis rotundis, apice acutis, base rotundatis, 1.5–3 cm
The type specimen of Capparis aegyptia has one longis, 1.2–2.5 cm latis, a C. zoharyi differt; stipulis
flower, no fruits. It is part of the herbarium A. T. tenuioribus, apicibus foliorum acutis nunquam rotundis nec
cordatis a C. zoharyi et C. aegyptia differt.
Danty d’Isnard, which is cited by Lamarck (1783):
‘‘v(idit) s(iccam) in herb(ario) Isn(ard).’’ It was Shrub procumbent, up to 60 cm high, glabrous;
included in 1857 in the Jussieu herbarium (P-JU). twigs straight, up to 2 m long, reddish purple,
The specimen was collected ‘‘en face de Minia’’ in sometimes green; internodes 1–3 cm; stipules curved,
Egypt by one botanist who gave it to Isnard: ‘‘Doni per retrorse, slightly decurrent, orange, 0.3–0.6 cm long,
tu’’ [Gift for you]. It was, presumably, D. Lippi (1678– 0.2–0.3 cm wide at the base. Leaves rounded, 1.5–3
1704) himself. Isnard wrote the manuscript list of 3 1.2–2.5 cm, somewhat fleshy; leaf veins not
plants—now at P-JU—collected by Lippi in Egypt prominent; bases rounded to cordate, apices acute;
during his last trip in 1704 (Pritzel, 1872). mucro very small, 0.1–0.5 mm, straight; petioles
The basionym of Capparis deserti (Zohary) Ta ¨ckh. & short, 0.3–1 cm. Flower buds rounded; floral pedicels
Boulos, Capparis spinosa L. var. deserti Zohary, was slender, short, 2–3 cm; flowers slightly zygomorphic;
validly published by Zohary. However he states ‘‘This abaxial (odd) sepal slightly galeate, 1.4–1.6 cm long,
is a ‘weak’ variety because small-leaved forms occur 0.5–0.6 cm deep; stamens 30 to 80, anthers very
almost in all groups.’’ small, 1.8–2 mm, apices rounded. Fruit oblong, pulp