Page 13 - CAPPARIS_2006

134 Annals of the
Missouri Botanical Garden

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Figure 5. Distribution map for Capparis parviflora ( ) (three subspecies); Capparis sicula ( ) (all five subspecies); and
*
Capparis mucronifolia (X) (both subspecies).
oblong-lanceolate, pubescence lax; flower buds United Arab Emirates]. Rocky slopes, ravines and
rounded ..... b. C. mucronifolia subsp. rosanoviana stony plains of deserts, wadies in Acacia seyal Del.
hammada, at elevations from 0 to 1000 m.
Distribution. Figure 5.
There is no designation of holotype for Capparis
4a. Capparis mucronifolia Boiss. subsp. mucro- mucronifolia by Boissier; the type material cited
nifolia comprises different specimens collected by P. M. R.
Aucher Eloy in southern Iran and Oman (former
Capparis elliptica Hausskn. & Bornm. ex Bornm. var.
maskatensis Hausskn. & Bornm. ex Bornm., Mitt. Thu ¨r. kingdom of Muscat), with collection numbers 4189,
Bot. Ver. N.F. VI: 49. 1894. TYPE: [Oman] ‘‘foliis 4190, and 4192, during his travels in 1835–1838.
latioribus ovatis; in rupibus ad Maskat, [J. Bornmu ¨ller] Therefore lectotypification is needed. The sheet with
ex. 46’’ (lectotype, designated here, JE!). the lectotype contains the following labels: I ‘‘Cap-
Shrub somewhat erect, heavily branched, irregu- paris mucronifolia / Boissier’’. II ‘‘Aucher-Eloy-
larly and widely spreading, up to 1 m high; twigs Herbier d’Orient Nu. 4189.’’ Paratypes are Aucher
slightly tortuose, approximately 2 m long, yellowish or pl. exs. 4190 and 4192, G!. Isolectoypes are in K!
grayish green; internodes 0.5–3 cm; stipules curved, Capparis elliptica Hausskn. & Bornm. ex Bornm.
retrorse, not decurrent, golden yellow, apex orange, Mitt. Thu ¨r. Bot. Ver. N.F. VI: 49. 1894, is a later
sometimes pubescent, at least basally, 0.2–0.6 cm homonym of Capparis elliptica Span. ex F. Muell.
long, 0.1–0.2 cm wide at the base. Leaves ovate, 2–4 Fragmenta Phytographiae Australiae 9: 172. 1875; it
3 0.5–1.5 cm, somewhat fleshy; indument very lax, is within the range of Capparis mucronifolia subsp.
trichomes thick and short to long, (20)30–40 3 200– mucronifolia.
400 mm; leaf veins not prominent; bases usually There is no designation of holotype for Capparis
rounded, sometimes tapering, apices acute; mucro elliptica var maskatensis Hausskn. & Bornm.; the type
small, 0.5–1.0 mm, straight; petioles very short, 0.2– material cited comprises different specimens collect-
0.4 cm. Flower buds acute; floral pedicels slender, ed, under number 46, by J. Bornmu ¨ller in his ‘‘Iter
short, 2.5–3.5 cm; flowers slightly zygomorphic; Persico–turcicum 1892–1893.’’ Haussknecht’s her-
abaxial (odd) sepal slightly galeate, 1.4–1.6 cm long, barium is at JE; Bornmu ¨ller worked from 1904 as
curator in JE, but sold his herbarium to B. Most of the
0.7–0.9 cm deep; stamens 30 to 80, anthers 1.3–
Bornmu ¨ller Capparidaceae material at B was de-
1.5 mm, with round apices. Fruit oblong, pulp color
stroyed during the Second World War. One Capparis
unknown; ripe seeds brown, 2.4–2.8 3 1.6–2 3 1.6–
specimen with collection number 46 is at JE (J.
1.8 mm.
Mu ¨ller, pers. comm.). Therefore lectotypification is
Phenology. Flowering from March to September. needed and possible. The sheet with the lectotype
Distribution and habitat. Sudano-Zambezian and contains the following labels: I ‘‘Isotypus/ Capparis
Saharo-Arabian, extending to the Irano-Turanian elliptica Hausskn. & Bornm./ var. maskatensis
Regions. Middle East [Afghanistan, Iran, Oman, Hausskn. & Bornm.’’ II ‘‘J. Bornmu ¨ller: Iter Persico-