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        314                              C. Brugnano et al. / Journal of Marine Systems 81 (2010) 312–322






































                                            Fig. 1. Study area and locations of sampling stations.

        every 50 m intervals from 100 to 300 m. Copepod species spatial  From 50 to 200 m depth (layers B, C, D, E), trends of decreasing
        distribution patterns were related to the environmental factors  temperature and increasing salinity with depth were registered.
        (dissolved oxygen, fluorescence, temperature and salinity) using  These waters which are probably a mixture of MAW and LIW, exhibit
        BIO-ENV procedure by superimposing the physico-chemical data on  temperature in the range of 14.34–16.27 °C and salinity in the range of
        the biotic ordination (Clarke and Warwick, 2004).    37.74–38.50.
                                                                Layer F (200–300 m depth, averages 14.28 °C, 38.74) represents
                                                             the Eastern Outflow Waters (EOW), i.e. eastern-origin waters already
        3. Results                                           undergone a partial mixing and layer G (300–600 m depth, averages
                                                             14.17 °C, 38.77) is the core LIW in the study area.
        3.1. Environmental parameters                           In the western-most stations (i.e. 7, 8, 9, 10, 11, 12 stations) MAW
                                                             influx is associated to a lower surface temperature than Tyrrhenian-
          The collected θ–Sprofiles (Fig. 2) show the hydrographical
        structure in the study area, which can be seen as a crossroad of
        eastern and western waters in the Mediterranean. The LIW signature
        with a salinity core of 38.77–38.78 was found from 300 m depth to the
        bottom in stations 10 and 12. After a partial mixing with resident
        waters the upper LIW vein enters the Tyrrhenian Sea and was present
        in the study area in the layer 200–300 m at stations 7, 10, 11, 12 with a
        core salinity of 38.72–38.74. In station 10, a subsurface core with
        salinity less than 37.5 in the layer 25–45 m (and a minimum of 37.29
        at 35 m) can be assigned to MAW. A similar feature was found at
        stations 1, 6 and 12 whereas, at stations 7, 8, 9 and 11 salinity always
        exceeded 37.5. So, the presence of MAW influences the upper 50 m-
        thick layer in the study area. To better finalise the use of
        hydrographical information to the objectives of the study, physico-
        chemical parameters of the whole data set were grouped by depth
        range according to the BIONESS vertical scheme of sampling and
        plankton data analysis. Vertical trends of temperature, salinity,
        dissolved oxygen and fluorescence are shown in Fig. 3. A clear
        thermocline is localized in the layer A* (20–40 m depth) with
        temperatures ranging from 23.5 to 16.2 °C. Higher temperatures and
        salinity in surface waters (averages in layer A: 22.85 °C, 37.90) are
        typical of a late summer condition whereas the subsurface minimum
        of salinity which can be noted (average in layer A*: 37.72) is the effect  Fig. 2. θ–S relationship for the collected CTD profiles around the Egadi Archipelago
        of fresh MAW influx in the region.                    (Stations 1 to 12, October 2004).
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